上嘴唇发白是因为什么原因

Gabunia L., Jahrb. RGZM 39, 185 (1992).

___, A.Vekua, Dmanissian Fossil Man and Accompanying Vertebrate Fauna (Metsniereba, Tbilisi, Georgia, 1993), pp. 1–71.

___, L'Anthropologie 99, 29 (1995).

Gabunia L., et al., Archa?ol. Korrespond. 29, 451 (1999).

Gabunia L., et al., Science 288, 1019 (2000).

L. Gabunia et al., in Early Humans at the Gates of Europe, D. Lordkipanidze, O. Bar-Yosef, M. Otte, Eds. (ERAUL 92, Liege, Belgium, 2000), pp. 13–27.

Lordkipanidze D., Vekua A., J. Hum. Evol. 42, A20 (2002).

Gabunia L., et al., Evol. Anthropol. 10, 158 (2001).

Gabunia L., Vekua A., Lordkipanidze D., J. Hum. Evol. 38, 785 (2000).

Behrensmeyer A. K., Paleobiology 2, 150 (1978).

M. Tappen et al., in Current Topics on Taphonomy and Fossilization, de Renzi et al., Eds. (Ajuntament de Valencia, Valencia, Spain, 2002), pp. 161–170.

Mimomys aff. pliocaenicus and Tchardinomys sp. from Dmanisi appear very close to M. pliocaenicus and Tchardinomys tegelensis from Tegelen discovered in sediments correlated with the upper part of the Olduvai Subchron (29, 30).

Justus A., Nioradze M., Mitt. Berl. Gesel. zur Anthrop. Ethn. Urg. Stuttgart 21, 61 (2001).

A. Walker, R. Leakey, The Nariokotome Homo erectus Skeleton (Harvard Univ. Press, Cambridge, MA, 1993), pp. 1–457.

Dean C., et al., Nature 414, 628 (2001).

McCollum M., et al. J. Hum. Evol 24 1993 87 .

Weidenreich F., Paleontol. Sinica New Ser. D 10, 1 (1943).

Gabunia L., Vekua A., Lordkipanidze D., Science 289, 55 (2000).

We elect to group early African fossils (also called H. ergaster) with H. erectus [sometimes H. erectus (sensu stricto)] as known from the Far East. B. Asfaw et al. (31) report recent finds from Bouri in Ethiopia demonstrating that there is continuity in morphology between the paleodemes of H. ergaster in East Africa and H. erectus in Asia. This evidence suggests that all the hominids may be treated as one polytypic species.

G. P. Rightmire, The Evolution of Homo erectus. Comparative Anatomical Studies of an Extinct Human Species (Cambridge Univ. Press, Cambridge, UK, 1990), pp. 1–260.

___, Am. J. Phys. Anthropol. 106, 61 (1998).

Gabunia L., Vekua A., Lordkipanidze D., Archeol. Ethnol. Anthropol. Eurasia 2, 128 (2001).

The large jaw (D2600) found in 2000 is high at the symphysis and has a long and relatively narrow alveolar arcade. The incisors (especially the I1s) are rather small-crowned. The canines are large but worn flat, with strong roots enclosed in massive juga. This specimen differs from D211 both in its dimensions and in the detailed morphology of the corpus, ascending ramus, and teeth. The index of robusticity is reduced as a result of great corpus height, shelving of the posterior face of the symphysis extends to the level of P4, canine juga are more pronounced, premolars are double-rooted, and the molars are larger, increasing slightly in size from M1 to M3 (see Table S1) (32).

Wood B., Nature 355, 783 (1992).

Several authors have argued that H. habilis (sensu stricto) and/or H. rudolfensis should be removed from Homo and placed instead with Australopithecus. J. T. Robinson (33) suggested this, and A. Walker (34) pointed out that the KNM-ER 1470 cranium exhibits a number of resemblances to Australopithecus. Recently, this view has been advanced by M. H. Wolpoff (35) and B. Wood and M. Collard (36).

P. V. Tobias, Olduvai Gorge, vol. 4, The Skulls, Endocasts and Teeth of Homo habilis (Cambridge Univ. Press, Cambridge, UK, 1991), pp. 1–921.

B. Wood, Koobi Fora Research Project, vol. 4, Hominid Cranial Remains (Clarendon, Oxford, UK, 1991).

Kimbel W. H., et al., J. Hum. Evol. 31, 549 (1996).

Zagwijn W. H., Mededeling. Nederl. Ins. Toegepast. Geowetensch. TNO 60, 19 (1998).

Tesakov A. S., Mededeling. Nederl. Ins. Toegepast. Geowetensch. TNO 60, 71 (1998).

Asfaw B., et al., Nature 416, 317 (2002).

L. L. Gabunia, M. A. De Lumley, A. Vekua, D. Lordkipanidze, C. R. Acad. Sci., in preparation.

Robinson J. T., Nature 205, 121 (1965).

A. Walker, in Earliest Man and Environments in the Lake Rudolf Basin, Y. Coppens, F. C. Howell, G. Ll. Isaac, R. F. Leakey, Eds. (Univ. of Chicago Press, Chicago, IL, 1976), pp. 484–489.

M. H. Wolpoff, Paleoanthropology (McGraw-Hill, New York, ed. 2, 1999).

Wood B., Collard M., Science 284, 65 (1999).

Research at Dmanisi is funded by the Georgian Academy of Sciences (grant N1318), National Geographic Society, and The Leakey Foundation (grants awarded to D.L.). Aspects of our interdisciplinary studies have been supported by Fulbright Foundation, Projects DGICYT-PB97-0157 (Spanish Ministry of Science) and ACE-38 (Generalitat de Catalunya), University of Zurich, the Eckler Fund of Binghamton University and the American School of Prehistoric Research, and the Peabody Museum of Harvard University. We thank all members of the 2001 Dmanisi research expedition, particularly J. Kopaliani, G. Kiladze, M. Mayer, G. Nioradze, S. Ediberidze, T. Shelia, D. Taktakishvili, and D. Zhvania, We are grateful to O. Bar-Yosef, F. C. Howell, H. de Lumley, M. A. de Lumley, and A. Walker for their help and assistance. Our work benefited from discussions with E. Delson, D. Lieberman, A. Justus, D. Pilbeam, O. Soffer, I. Tattersall, M. Wolpoff, and B. Wood. CT scans were produced at the Medical-Diagnostic Center of Tbilisi University. Photographs and illustrations were made by G. Davtiani, S. Holland, and G. Tsibakhashvili. Supporting Online Material www.sciencemag.org/content/full/297/5578/85/DC1 Table S1 SFigs. 1 and S2